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After speaking with your faction's representative in your respective pavillion you will be sent on several quests that teach you how to use the jousting combat system:
Mastery of Melee - The first thing you need to do with this quest is equip the lance you are given. Then, go outside the tent and jump on the vehicle horse or hawkstrider. It then sends you to talk to a training NPC that will teach you how to use the Defend and Thrust abilities. You need to practice these on the targeting dummies. Dummies for all quests are located in the north area of the Argent Tournament, above the coliseum-in-progress. The melee target dummy is farthest west. Prior to approaching the dummy use your Defend ability three times to get maximum protection. Floating shields in red (1), yellow (2), and green (3) indicate your Defend stack. Approach the targeting dummy and attack with button 1: Thrust. The dummy will hit you back, removing a charge from your Defend, so you need to keep using the ability to reinforce your defenses. You must successfully use Thrust on the targeting dummy five times to complete the quest.
Mastery of the Shield Breaker - As with the above, equip your lance, jump on the mount, and go to the area with the targeting dummies. Speak to the designated NPC to learn more about the ability. You need to use the Shield-breaker ability while standing away from the Ranged Targets. You'll need to use it five times: 3 to break through the defenses, and twice to hit the target for quest credit.
Mastery of the Charge - Same as above, but this time you'll be beating up on the Charge Target dummies. Use Shield-breaker before charging the dummy. When you charge your vehicle will continue running a good distance past the target - don't be alarmed. Do this twice to complete the quest.